The Republic of Georgia. East Georgia; Mtkheta district; Saguramo range; Saguramo reservation, on the road from vill. Saguramo to Zedazeni, ca. 41°52'39N, 44°46'26E, ca. 1000 m. (coordinates from hand-held GPS, accurate to within a radius of about 50 m).
Republic of Georgia : Lagodekhi Nature Reserve. Along the trail above timberline southeast of the meterological station (''Meteo Sta'') and field cabin from timberline to a minor pas on the trail from Meteo Station to Lekisaplavi.
Reveal, J.|Mamukelashvili Gogi, Temur Siukaev.