Antsiranana, Ankarana special reserve, 6 km NE Mahatsara village, sur la route Mahatsara-Diego Suarez road, near volcanic outcrops, in pasture, in primary forest patch
Toamasina, Phelps Dodge project site, ca. 15 air-km NE of Moramanga, ca. 11 km E of Antanambao. Between Ambatovy-South and Analamay-East, Torotorofotsy River, Berano hillside gradient
Rakotomalaza, P.J.|Razafimanantsoa, G.|Andriatsiferana, F.
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