E: Toamasina (Tamatave) Province, c. 2 km East of Ampasimanolotra (Brickaville), near the start of the road to Ambila-Lemaitso. (Also seen on Nosy Mangabe)
SC: Fianarantsoa Province, 29 km NW of Ranohira, on the eastern edge of the south-eastern corner of the Isalo National Park, ruiniform sandstone massif
SW: Toliara (Tuléar) Province, E of Betioky, Valley of the Salohilova River, offshoot of the Hazoara River, leading into the Sakamena River, c. 6 km south of Parcelle 1 of Beza Mahafaly Reserve
Du Puy, D.J.|Du Puy, B.P.|Ravonjiarisoa, P.|Phillipson, P.B.
M 73
SW: Toliara (Tuléar) Province, E of Betioky, Valley of the Salohilova River, offshoot of the Hazoara River, leading into the Sakamena River, c. 6 km south of Parcelle 1 of Beza Mahafaly Reserve
Du Puy, D.J.|Du Puy, B.P.|Ravonjiarisoa, P.|Phillipson, P.B.
M 74
SW: Toliara (Tuléar) Province, E of Betioky, Valley of the Salohilova River, offshoot of the Hazoara River, leading into the Sakamena River, c. 6 km south of Parcelle 1 of Beza Mahafaly Reserve
Du Puy, D.J.|Du Puy, B.P.|Ravonjiarisoa, P.|Phillipson, P.B.
M 75
SW: Toliara (Tulear) Province, small track towards Tsefanoka, from near Tsianisiha (on Route Nationale 9), valley of Manombo River, c. 60 km N of Toliara
SW: Toliara (Tulear) Province, small track towards Tsefanoka, from near Tsianisiha (on Route Nationale 9), valley of Manombo River, c. 60 km N of Toliara
Fianarantsoa Province, Vatovavy-Fitovinany Region, Ifanadiana District, Ranomafana National Park, along the main trail from the main entrance of Vohipara
Expedition 2016 to Madagascar of the Botany Department of the Swedish Museum of Natutal History 1653
Swenson, U.|Razafindrahaja, V.|Atalahy, A.|Razafimandimbison, S.
1653