Toliara : Andohahela RNI, parcelle 2, c. 50 km WNW (BRG 295°) of Taolanaro, riverine/galley forest (tributary of Mananara river), c. 5 km due E of Ihazofotsy, on the banks of the Mananara.
Toamasina : Phelps Dodge project site, ca. 15 air-km NE of Moramanga, ca. 11 km E of Antanambao. Ambatovy-South, wetland south (north-east branch), hillside gradient