Antsiranana. Réserve naturelle Intégrale de Marojejy, 11 km NW of Manantenina, forest above "Camp 3" (where GPS reading was taken) along trail to summit
Fianarantsoa, along National road 25 W of Ranomafana ; between the towns of Ranomafana and Ambatolahy. Found ca. 300 m downsteam from JIRAMA power station, ca. 500 m S of National road 25