Toamasina : Masoala Peninsula, south of the village of Ambanizana in the Andranobe River Watershed, Between 400 and 1700 m from the seashore, 90 degree a,d 9 m from trail point K100
Andapa, Doany, Betsomanga, environ à 11.2 km à vol d'oiseau au SE de Doany et à 7.5 km à vol d'oiseau au NE d'Ambalamanasy II, entre la rivière d'Ampandrana et Betsomanga, au N de Marojejy
Antsiranana, Montagne d'Ambre National Park. ca. 1,5 km E of intersection of main road to forestry station and road to Grande Cascade, from primary forest and from disturbed area near Maison de Guardian de Station
Toliara : Préfecture de Tôlanaro (Fort-Dauphin); foot trail beyond end of track on N side of Antorendrika River ; ca 22 km NE of Tolanaro (Fort-Dauphin)