Antsiranana. Réserve Naturelle de Marojejy, along the trail to the summit of Marojejy Est, N of Mandena ; lowland rainforest between the southern limit of the reserve and the lst camp.
Toamasina. Masoala Peninsula, south of the village of Ambanizana in the Andranobe River Watershed. Betwenn 400 and 1700 m from the seashore in primary lowland rain forest