Reserve Naturelle de Raleigh Falls (Riv. Coppename), inselbergs du Voltzberg, savane roche sur l'affleurement rocheux au pied du premier piton du Voltzberg, près du camp
Depto. Santa Cruz, Prov. Vallegrande, steep slopes of valley of the rio Grande, on road from Pucará to the new bridge over the rio Grande, 16km. (by winding road) NW of Puente Santa Rosa. Bolívia
Apure, distrito Pedra Camejo, banks of the Rio Orinoco on Isla Poyatón opposite the Serrania de Baraguán on the Apure side of Estrecho de Baraguán of the Rio Orinoco. Venezuela
Miranda: Distrito Páez, near the upper limit of the flood basin of the El Guapo Repressa below the junction of the Río Guapo, ca. 11km SW. (in a straight line) of El Guapo; forested canyon bottom along the Rìo Guapo. Venezuela