Antsiranana: Masoala Peninsula, forest surrounding research station at Andronabe on western coast.Two altitudinal transects each with stations 1-7 at altitudinal increments from 0 m to 600 m altitude respectively
Fianarantsoa: Andringitra Reserve N°. 5, S of Ambalavao; open, level areas in cirque S and above village of Antanifotsy; partially disturbed, probably burned areas dominated by grasses