Toliara : Préfecture de Tôlagnaro (Fort-Dauphin), Canton de Manambaro, Petriky forest, due N of NE corner of Lake Andranany and NW of large dune at end of road, in low forest on gray sand (ca. 11 km WSW of Tôlagnaro)
Fianarantsoa : Sud-Est de Fianarantsoa, à l'extérieur du Parc National de Ranomafana, Parcelle III. A 25 km au Sud-Ouest de Ranomafana. Autours de Sahavanana
Toliara : Fort-Dauphin (Tôlagnaro) region ; NE of city, in region called Mandena, beyond Botanical Garden and QIT camp, in forest remnants near coastal lake
Toamasina : Phelps Dodge project site, ca. 15 air-KM NE of Moramanga, ca. 11 km E of Antanambao. Ambatovy, Antsahalava River east, hillside gradient : zonal mid-altitude rain forest, selectively logged (pump-station)