Antsiranana. Masoala peninsula, forest surrounding research station at Andronabe on western coast. Two altitudinal transects each with stations 1-7 at altitudinal increments
Joffreville Distr.: Ca. 16.4 km SW of main road between Diego Suarez and on Joffreville Road to P.N. Montagne D'Ambre and ca. 4 km E of Joffreville town center. In primary forest patch ca. 500 m from road
Toamasina : Phelps Dodge project site, ca. 15 air-km NE of Moramanga, ca. 11 km E of Antanambao. Ambatovy-South, wetland south (north-east branch), hillside gradient
Antsiranana : Environs de la Réserve Spéciale d'Ankarana, à 20 km au Nord d'Ambilobe. A 4 km à l'Ouest du village de Betsimiranjana, Campement près du village d'Ambiba