Mpio Rangel. Paramo de Motumbo, SW of Laguna Larga (Trujillo state) along border of divide to Laguna Las Parias (Trujillo state), Monumento Natural Teta de Niquitao-Guirigay.
Province Zamora-Chinchipe : Miazi, at junction of Rio Chumbiriatza with Rio Nangaritza. Tall montane forest, riverside forest, riverbanks, dwarfted mossy ridgetop forest and calcareous rocks