S Madagascar, province of Toliara (Tulear) Reserve naturelle Integrale 11, Andohahela (parcelle 1), path to Iminiminy, col de Antanatana, before Andranomitidy
Du Puy, D.J.|Du Puy, B.P.|Cheek, M.|Rafamantanantsoa, G.
M 398
Ranomafama national park, vicinity of Ampasina, a circa 150 ha flat marshy area located 3-4 hours walk NE Amboditanimena (nat. road between Vohiparara and Ambalakindresy)