Province Diego-Suarez: forêt de Mahavinitra, Réserve Naturelle de Masoala, vicinity of Ambatobé, accross the Onive river and about two hours by foot upriver from Ambohitralanana, Antalaha Prefecture.
Est : Forêt d'Ankiririryra east of the village of Ambuddiapaly, one hour by foot from the manandrana River at crossing of road from Antaratasy between Tamatave and Foulpointe