Toamasina Prov., near Antandroroho village, 16 km West of Mahanoro, on unconsolidated sand bordering the Amparafana littoral forest under three co-dominating tree species, viz. Uapaca littoralis, U. thouarsii ( Phyllantaceae) and Leptolaena multiflora (Sarcolaenaceae)
East Province, Dja Biosphere Reserve, Northwest Sector near the village
of Somalomo, Upper Dja River Basin, within 2 km radius of Dja base camp located at 3° 21' 29.8"
N; 12° 43' 46.9" W