Bassin de réception de la Mananara, affluent du Mandrare ; pentes occidentales des montagnes entre l'Andohahela et l'Elakelaka, entre Ampahiso et Mahamavo
Toliara : 55 km NE of Morondava via route 8 at the forestry concession of the Centre de Formation Professional Forestière (CFPF), Kirindy Forest. 5 km E of route 8 along piste CONOCO, alond S boundary of forestry bloc CN 5
Toamasina, Phelps dodge project site, ca. 15 air-km NE of Moramanga, ca. 11 km E of Antanambao. Between Ambatovy-South and Analamay-East, Torotorofotsy river, Berano