Fivondronana : Fort-Dauphin, Firaisana : Mahatalaky. Fokontany, Sainte Luce. Forêt littorale sur sables de Sainte Luce, bloc S9 de la zone de conservation QMM
Fivondronana, Tolagnaro, N of Tolagnaro, Mandena forestry station, littoral forest on sand, along the first main trail E-W, not far from the guardian house