Joculator Hedley, 1909

Zoology

Taxonomy

Species

Joculator acuminatus Cecalupo & Perugia, 2012 18 3

Joculator adustus Cecalupo & Perugia, 2024 1

Joculator aeneus Cecalupo & Perugia, 2017 2 2

Joculator albatus Cecalupo & Perugia, 2023 1

Joculator alligatus Cecalupo & Perugia, 2012 10 3

Joculator anconae Cecalupo & Perugia, 2017 1 1

Joculator angustus Cecalupo & Perugia, 2017 5 2

Joculator antonioi Cecalupo & Perugia, 2012 7 1

Joculator aoreensis Cecalupo & Perugia, 2013 1 1

Joculator arduinii Cecalupo & Perugia, 2012 7 2

Joculator arenaceus Cecalupo & Perugia, 2013 1 1

Joculator ariadnae Cecalupo & Perugia, 2018 1 1

Joculator asper Cecalupo & Perugia, 2024 1

Joculator ater Cecalupo & Perugia, 2012 2 1

Joculator autumnus Cecalupo & Perugia, 2016 1 1

Joculator bangkaensis Cecalupo & Perugia, 2019 1 1

Joculator baraggiai Cecalupo & Perugia, 2013 1 1

Joculator barteri Cecalupo & Perugia, 2018 2 2

Joculator bauquisi Cecalupo & Perugia, 2017 4 2

Joculator berberiani Cecalupo & Perugia, 2018 1 1

Joculator bicinctus Cecalupo & Perugia, 2012 26 3

Joculator boguszae Cecalupo & Perugia, 2018 2 1

Joculator bourgeoisae Cecalupo & Perugia, 2013 5 2

Joculator bouteti Cecalupo & Perugia, 2014 11 5

Joculator brabantae Cecalupo & Perugia, 2013 3 2

Joculator breviculus Cecalupo & Perugia, 2017 2 2

Joculator brevis Cecalupo & Perugia, 2012 3 1

Joculator brisseti Cecalupo & Perugia, 2021 1 1

Joculator brusonii Cecalupo & Perugia, 2013 3 1

Joculator caliginosus Cecalupo & Perugia, 2012 2 2

Joculator caloi Cecalupo & Perugia, 2013 3 2

Joculator carpatinus Cecalupo & Perugia, 2012 5 2

Joculator castaneus Cecalupo & Perugia, 2013 1 1

Joculator cereus Cecalupo & Perugia, 2012 3 1

Joculator cf. christiaensi Jay & Drivas, 2002 1

Joculator cf. dupouxae Cecalupo & Perugia, 2018 1

Joculator cf. recisus Cecalupo & Perugia, 2012 1

Joculator cf. saguili Cecalupo & Perugia, 2013 2

Joculator christiaensi Jay & Drivas, 2002 1 1

Joculator cinctus Cecalupo & Perugia, 2012 5

Joculator citrinus Cecalupo & Perugia, 2013 5 1

Joculator coffeus Cecalupo & Perugia, 2013 9 5

Joculator conformis Cecalupo & Perugia, 2013 1 1

Joculator corneliae Cecalupo & Perugia, 2018 1 1

Joculator cossignanii Cecalupo & Perugia, 2012 6 3

Joculator cristinae Cecalupo & Perugia, 2013 3 2

Joculator czubai Cecalupo & Perugia, 2018 1 1

Joculator dakoi Cecalupo & Perugia, 2018 3 1

Joculator desmieri Cecalupo & Perugia, 2018 2 1

Joculator diversus Cecalupo & Perugia, 2019 1 1

Joculator drivasi Cecalupo & Perugia, 2012 1 1

Joculator dupouxae Cecalupo & Perugia, 2018 1 1

Joculator emidioi Cecalupo & Perugia, 2014 2 1

Joculator erianae Cecalupo & Perugia, 2017 4 2

Joculator eudeli Jay & Drivas, 2002 3

Joculator faninozi Cecalupo & Perugia, 2018 6 1

Joculator fedosovi Cecalupo & Perugia, 2018 2 1

Joculator ferrii Cecalupo & Perugia, 2013 17 1

Joculator ferrugineus Cecalupo & Perugia, 2012 2 1

Joculator filum Cecalupo & Perugia, 2017 3 3

Joculator fischeri Jay & Drivas, 2002 2

Joculator flavicans Cecalupo & Perugia, 2014 1 1

Joculator frequens Cecalupo & Perugia, 2012 17 1

Joculator furvus Cecalupo & Perugia, 2012 2 2

Joculator fuscostriatus Cecalupo & Perugia, 2013 5 2

Joculator fuscus Cecalupo & Perugia, 2012 7 1

Joculator gagulae Cecalupo & Perugia, 2018 4 1

Joculator ganui Cecalupo & Perugia, 2018 3 1

Joculator garianii Cecalupo & Perugia, 2013 2 2

Joculator gemmae Cecalupo & Perugia, 2012 3 2

Joculator giovanolii Cecalupo & Perugia, 2013 8 2

Joculator gorini Cecalupo & Perugia, 2014 5 1

Joculator goubini Cecalupo & Perugia, 2017 2 1

Joculator granulosus Cecalupo & Perugia, 2013 3 1

Joculator grimaldae Cecalupo & Perugia, 2021 1 1

Joculator hadfieldae Cecalupo & Perugia, 2017 3 3

Joculator herosae Cecalupo & Perugia, 2012 1 1

Joculator humilis Cecalupo & Perugia, 2012 2 2

Joculator ianthinus Cecalupo & Perugia, 2013 3 1

Joculator incisus Cecalupo & Perugia, 2012 5 1

Joculator inflatus Cecalupo & Perugia, 2012 1 1

Joculator iohannae Cecalupo & Perugia, 2012 3 1

Joculator iopuei Cecalupo & Perugia, 2017 6 4

Joculator itiensis Cecalupo & Perugia, 2014 21 1

Joculator jeffkinchi Cecalupo & Perugia, 2018 1 1

Joculator johnkasui Cecalupo & Perugia, 2018 1 1

Joculator juliuschani Cecalupo & Perugia, 2018 1 1

Joculator kaipuae Cecalupo & Perugia, 2018 3 3

Joculator kanoi Cecalupo & Perugia, 2013 4 4

Joculator kenganziki Cecalupo & Perugia, 2018 1 1

Joculator keratochroma Jay & Drivas, 2002 2

Joculator keszlerae Cecalupo & Perugia, 2021 2 1

Joculator koluae Cecalupo & Perugia, 2021 1 1

Joculator kuboi Cecalupo & Perugia, 2019 1

Joculator laguncula (Cecalupo & Perugia, 2012) 1 1

Joculator laregnereensis Cecalupo & Perugia, 2017 2 2

Joculator laseroni Jay & Drivas, 2002 2

Joculator latrechei Cecalupo & Perugia, 2021 1 1

Joculator lazzarii Cecalupo & Perugia, 2013 4 3

Joculator legallae Cecalupo & Perugia, 2014 2 1

Joculator legoffi Cecalupo & Perugia, 2017 2 1

Joculator leguyaderi Cecalupo & Perugia, 2013 3 2

Joculator lividus Cecalupo & Perugia, 2012 16 5

Joculator lokaroensis Cecalupo & Perugia, 2014 1 1

Joculator lozoueti Jay & Drivas, 2002 2

Joculator luteolus Cecalupo & Perugia, 2012 1 1

Joculator maestratii Lozouet, 2001 2

Joculator magninensis Cecalupo & Perugia, 2017 1 1

Joculator maloensis Cecalupo & Perugia, 2013 3 1

Joculator manneensis Cecalupo & Perugia, 2018 1 1

Joculator maranii Cecalupo & Perugia, 2014 2 1

Joculator marinae Cecalupo & Perugia, 2014 2 1

Joculator massimilianoi Cecalupo & Perugia, 2012 1 1

Joculator mauberti Cecalupo & Perugia, 2018 1 1

Joculator mbereensis Cecalupo & Perugia, 2017 1 1

Joculator meanii Cecalupo & Perugia, 2014 1 1

Joculator megacephala Jay & Drivas, 2002 2

Joculator melanoraphis Jay & Drivas, 2002 2

Joculator micalii Cecalupo & Perugia, 2012 5 1

Joculator michoni Cecalupo & Perugia, 2018 2 1

Joculator minimului Cecalupo & Perugia, 2018 3 1

Joculator minimus Laseron, 1956 2 2

Joculator minutissimus (Thiele, 1925) 1

Joculator minutus Cecalupo & Perugia, 2012 7 2

Joculator mitsuoi Cecalupo & Perugia, 2019 1

Joculator modestus Cecalupo & Perugia, 2012 4 2

Joculator morenae Cecalupo & Perugia, 2013 1 1

Joculator morigii Cecalupo & Perugia, 2017 2 1

Joculator murciai Cecalupo & Perugia, 2014 1 1

Joculator mygaki Jay & Drivas, 2002 4 1

Joculator myia Jay & Drivas, 2002 2

Joculator nitidus Cecalupo & Perugia, 2012 1 1

Joculator nucleus Cecalupo & Perugia, 2019 1 1

Joculator obscurus Cecalupo & Perugia, 2012 1 1

Joculator obsoletus Cecalupo & Perugia, 2012 1 1

Joculator occultus Cecalupo & Perugia, 2012 3 2

Joculator olivoideus Cecalupo & Perugia, 2018 1 1

Joculator onnaensis Cecalupo & Perugia, 2019 1

Joculator opulentus Cecalupo & Perugia, 2013 4 2

Joculator pallidus Cecalupo & Perugia, 2012 2 1

Joculator parvulus Cecalupo & Perugia, 2012 10 2

Joculator pauxillus Cecalupo & Perugia, 2012 1 1

Joculator perforatus Cecalupo & Perugia, 2013 2 1

Joculator perlucidus Cecalupo & Perugia, 2012 1 1

Joculator phtyr Jay & Drivas, 2002 2

Joculator pierresamueli Cecalupo & Perugia, 2021 2 2

Joculator pinguis Cecalupo & Perugia, 2012 1 1

Joculator pizzinii Cecalupo & Perugia, 2013 4 2

Joculator priorai Cecalupo & Perugia, 2012 4 2

Joculator problematicus Albano & Steger, 2021 1 1

Joculator prunus Cecalupo & Perugia, 2013 3 1

Joculator psyllos Jay & Drivas, 2002 2

Joculator pumilus Cecalupo & Perugia, 2018 3 2

Joculator pupiformis Cecalupo & Perugia, 2012 6 1

Joculator pygmaeus Cecalupo & Perugia, 2012 3 2

Joculator quaggiottoi Cecalupo & Perugia, 2012 4 1

Joculator ralijaonai Cecalupo & Perugia, 2014 2 1

Joculator ralphmanai Cecalupo & Perugia, 2018 1 1

Joculator rambelli Cecalupo & Perugia, 2018 3 1

Joculator recisus Cecalupo & Perugia, 2012 3 2

Joculator rolani Cecalupo & Perugia, 2012 2 1

Joculator rubus Cecalupo & Perugia, 2013 2 2

Joculator sabrinae Cecalupo & Perugia, 2012 4 1

Joculator saguili Cecalupo & Perugia, 2013 7 3

Joculator salvati Jay & Drivas, 2002 2

Joculator sarahpezetae Cecalupo & Perugia, 2018 3 2

Joculator savellii Cecalupo & Perugia, 2021 1 1

Joculator schiaparellii Cecalupo & Perugia, 2017 3 2

Joculator sekensis Cecalupo & Perugia, 2018 1 1

Joculator semiperlucidus Cecalupo & Perugia, 2013 2 2

Joculator simonsili Cecalupo & Perugia, 2018 4 1

Joculator simulans Cecalupo & Perugia, 2012 4 3

Joculator siwisikae Cecalupo & Perugia, 2018 2 1

Joculator skolix Jay & Drivas, 2002 2

Joculator sp. 17

Joculator steykerae Cecalupo & Perugia, 2013 3 2

Joculator stramineus Cecalupo & Perugia, 2013 1 1

Joculator subconicus Cecalupo & Perugia, 2012 4 2

Joculator subdolus Cecalupo & Perugia, 2012 2 2

Joculator subglobosus Cecalupo & Perugia, 2013 9 6

Joculator succineus Cecalupo & Perugia, 2013 3 1

Joculator sylviakinchae Cecalupo & Perugia, 2018 1 1

Joculator tagliaferroi Cecalupo & Perugia, 2017 2 2

Joculator testaceus Cecalupo & Perugia, 2013 2 1

Joculator testii Cecalupo & Perugia, 2017 2 2

Joculator theresae Cecalupo & Perugia, 2018 4 1

Joculator thielei Jay & Drivas, 2002 1

Joculator tieensis Cecalupo & Perugia, 2017 3 1

Joculator trifasciatus Cecalupo & Perugia, 2024 1

Joculator tsiriveloi Cecalupo & Perugia, 2014 3 1

Joculator unicolor Cecalupo & Perugia, 2012 4 3

Joculator vandelae Cecalupo & Perugia, 2014 1 1

Joculator variabilis Cecalupo & Perugia, 2012 7 3

Joculator vassardi Cecalupo & Perugia, 2014 2 1

Joculator vazzanai Cecalupo & Perugia, 2018 3 1

Joculator venustus Cecalupo & Perugia, 2013 3 1

Joculator vignali Jay & Drivas, 2002 3

Joculator violaceus Cecalupo & Perugia, 2012 5 2

Joculator voncoseli Cecalupo & Perugia, 2012 7 2

Joculator websterae Cecalupo & Perugia, 2014 1 1

Joculator zahariasi Cecalupo & Perugia, 2021 1 1

Joculator ziliolii Cecalupo & Perugia, 2012 6 2

Joculator zucconi Cecalupo & Perugia, 2021 1 1

Specimens registered in collections

Collection	Number of specimens (or sets)
Molluscs (IM)	654